• Goldsby, H., B. Kerr, & C. Ofria (2020) "Major transitions in digital evolution." "Evolution in Action: Past, Present and Future." Eds. W. Banzhaf et al. Springer International Publishing, 333-347. (Link)
  • Tanaka, M., P. Godfrey-Smith, & B. Kerr (2020) The dual landscape model of adaptation and niche construction. Philosophy of Science (Link)
  • Jordt, H., T. Stalder, O. Kosterlitz, J. M. Ponciano, E. M. Top & B. Kerr (2020) Coevolution of host–plasmid pairs facilitates the emergence of novel multidrug resistance. Nature Ecology & Evolution: (DOI) (PDF)
  • Stalder, T., B. Cornwell, J. Lacroix, B. Kohler, S. Dixon, H. Yano, B. Kerr, L. J. Forney, & E. M. Top (2020) Evolving populations in biofilms contain more persistent plasmids. Molecular Biology and Evolution: (DOI) (PDF)


  • Godfrey-Smith, P. & B. Kerr (2019) Tolerance: A hierarchical analysis. Journal of Political Philosophy 27: 403-421. (link)


  • Chisholm, R. H., B. D. Connelly, B. Kerr, M. M. Tanaka (2018) The role of pleiotropy in the evolutionary maintenance of positive niche construction. American Naturalist 192 (link)


  • Singhal, S., C. M. Leon Guerrero, S. G. Whang, E. M. McClure, H. G. Busch, B. Kerr (2017) Adaptations of an RNA virus to increasing thermal stress. PLoS ONE 12: (e0189602). (PDF)
  • Nahum, J. R., J. D. West, B. A. Althouse, L. Zaman, C. Ofria, and B. Kerr (2017) Improved adaptation in exogenously and endogenously changing environments. Proceedings of European Conference on Artificial Life (ECAL), 14: 306-313. (PDF)
  • Kokko, H., A. Chaturvedi, D. Croll, M. C. Fischer, F. Guillaume, S. Karrenberg, B. Kerr, G. Rolshausen, and J. Stapley. (2017) Can Evolution Supply What Ecology Demands? Trends in Ecology and Evolution 32: 187–197. (PDF)
  • Wargo, A. R., R. J. Scott, B. Kerr, and G. Kurath. (2017) Replication and shedding kinetics of infectious hematopoietic necrosis virus in juvenile rainbow trout. Virus Research 227: 200-211. (PDF)


  • Cooper, J. and B. Kerr (2016) Evolution at ‘sutures’ and ‘centers’: recombination can aid adaptation of spatially structured populations on rugged fitness landscapes. PLoS Computational Biology (PDF)
  • Libby*, E., P. L. Conlin*, B. Kerr, W. C. Ratcliff (2016) Stabilizing multicellularity through ratcheting. Phil. Trans. R. Soc. B 371: 20150444. (PDF)
  • van Raay, K. and B. Kerr (2016) Toxins go viral: phage-encoded lysis releases group B colicins. Environmental Microbiology 18: 1308–1311. (PDF)
  • Estrela, S., B. Kerr, and J. J. Morris (2016) Transitions in individuality through symbiosis. Current Opinion in Microbiology 31: 191–198. (PDF)
  • Hammarlund*, S. P., B. D. Connelly*, K. J. Dickinson, and B. Kerr (2016) The evolution of cooperation by the Hankshaw effect. Evolution 70: 1376–1385. (PDF)
  • Connelly, B. D., K. J. Dickinson, S. P. Hammarlund, and B. Kerr (2016) Negative niche construction favors the evolution of cooperation. Evolutionary Ecology 30:267–283. (PDF)
  • Estrela, S., J. J. Morris, and B. Kerr (2016) Private benefits and metabolic conflicts shape the emergence of microbial interdependencies. Environmental Microbiology 18, 1415–1427. (PDF)


  • Miner, B. E., P. M. Kulling, K. D. Beer, and B. Kerr (2015) Divergence in DNA photorepair efficiency among genotypes from contrasting UV radiation environments in nature. Molecular Ecology 24: 6177–6187. (PDF)
  • Heilmann, S., S. Krishna, and B. Kerr (2015) Why do bacteria regulate public goods by quorum sensing?—How the shapes of cost and benefit functions determine the form of optimal regulation. Frontiers in Microbiology 6: 1-11. (PDF)
  • Cooper, J. D., C. Neuhauser, A. M. Dean, and B. Kerr (2015) Tipping the mutation–selection balance: Limited migration increases the frequency of deleterious mutants. Journal of Theoretical Biology 380: 123-133. (PDF)
  • Bergstrom, C. and B. Kerr (2015) Microbiology: Taking the bad with the good. Nature (PDF) (Focal Article) (Supplemental Model)
  • Nahum, J. R., P. Godfrey-Smith, B. N. Harding, J. H. Marcus, J. Carlson-Stevermer, and B. Kerr (2015) A tortoise–hare pattern seen in adapting structured and unstructured populations suggests a rugged fitness landscape in bacteria. Proceedings of the National Academy of Science 112: 7530–7535. (PDF) (Commentary) (News Item)


  • Hammerschmidt, K., C. Rose, B. Kerr, P. Rainey (2014) Life cycles, fitness decoupling and the evolution of multicellularity. Nature 515: 75-79. (PDF) (News Item)
  • Conlin, P., J. Chandler, B. Kerr (2014) Games of life and death: antibiotic resistance and production through the lens of evolutionary game theory. Current Opinion in Microbiology 21: 35-44. (PDF)
  • Libby, E., W. Ratcliff, M. Travisano, B. Kerr (2014) Geometry shapes evolution of early multicellularity. PLoS Computational Biology 10(9): e1003803. doi:10.1371/journal.pcbi.1003803 (PDF) (News Item)
  • Goldsby HJ, Knoester DB, Kerr B, Ofria C (2014) The effect of conflicting pressures on the evolution of division of labor. PLoS ONE 9(8): e102713. doi:10.1371/journal.pone.0102713 (PDF)
  • Goldsby H.J., D.B. Knoester, C. Ofria and B. Kerr. 2014. The evolutionary origin of somatic cells under the dirty work hypothesis. PLoS Biology 12 (5): e1001858. doi:10.1371/journal.pbio.1001858 (PDF) (Dryad Data Deposit) (PLoS Synopsis) (News Item) (Blog Post by Sedeer el-Showk) (Blog Post at Biophilia)



  • Goldsby, H., A. Dornhaus, B. Kerr and C. Ofria. 2012. Task-switching costs promote the evolution of division of labor and shifts in individuality. Proceedings of the National Academy of Science 109: 13686–13691 (PDF)
  • Goldsby, H., T. Pierce, F. Dyer, B. Kerr and C. Ofria. 2012. The evolution of temporal polyethism. Artificial Life 13: 178-185 (PDF)
  • Fox, J. W. and B. Kerr. 2012. Analyzing the effects of species gain and loss on ecosystem function using the extended Price equation partition. Oikos 121: 290-298 (PDF)


  • Nahum, J. R., B. N. Harding and B. Kerr. 2011. Evolution of restraint in a structured rock-paper-scissors community. Proceedings of the National Academy of Science 108: 10831-10838 (PDF) (Supporting Information) (Wiki Supplement)
  • Kerr, B. and J. Nahum. 2011. Setting the stage for an egalitarian major transition: The evolution of restraint. For—The Major Transitions Revisited: The Evolution of Individuality Eds. K. Sterelny & B. Calcott. (Link)
  • Rainey, P. and B. Kerr. 2011. Conflicts among levels of selection as fuel for the evolution of individuality. For—The Major Transitions Revisited: The Evolution of Individuality Eds. K. Sterelny & B. Calcott. (Link)
  • Miner, B. and B. Kerr. 2011. Adaptation to local ultraviolet radiation conditions among neighbouring Daphnia populations. Proceedings of the Royal Society B 278:1306-1313 (PDF) (Supplement)
  • Majeed, H., O. Gillor, B. Kerr and M. Riley. 2011. Competitive interactions in Escherichia coli populations: the role of bacteriocins. The ISME Journal.  5: 71–81. (PDF


  • Eshelman, C., R. Vouk, J. Stewart, E. Halsne, H. Lindsey, S. Schneider, M. Gualu, A. Dean and B. Kerr. 2010. Unrestricted migration favours virulent pathogens in experimental metapopulations: Evolutionary genetics of a rapacious life history. Philosophical Transactions of the Royal Society B. 365: 2503-2513. (PDF) (Supplement)
  • Rainey, P. and B. Kerr. 2010. Cheats as first propagules: a new hypothesis for the evolution of individuality during the transition from single cells to multicellularity. Bioessays. 32: 872–880. (PDF)


  • Prado, F., T. Billo, and B. Kerr. 2009. Introgression of sexually selected traits in lek-mating species. Evolutionary Ecology Research 11: 1235-1250. (PDF)
  • Godfrey-Smith, P. and B. Kerr. 2009. Selection in ephemeral networks. American Naturalist 174: 906-911. (PDF)
  • Prado, F., A. Sheih, J. West, and B. Kerr. 2009. Coevolutionary cycling of host sociality and pathogen virulence in contact networks. Journal of Theoretical Biology 261: 561-569. (PDF)
  • Dunlap, A., C. McLinn, H. MacCormick, M. Scott, and B. Kerr. 2009. Why some memories don’t last a lifetime: Optimal long-term recall in changing environments. Behavioral Ecology 20: 1096-1105. (PDF)
  • Kerr, B. 2009. "Theoretical and experimental approaches to the evolution of altruism and the levels of selection." Experimental Evolution: Concepts, Methods, and Applications of Selection Experiments. Eds. T. Garland & M. R. Rose. Berkeley, CA: University of California Press, 585-630. (PDF)
  • Gobush, K., B. Kerr, and S. Wasser. 2009. Genetic relatedness and disrupted social structure in a poached population of African elephants. Molecular Ecology 18: 722-734. (PDF)
  • Kerr, B., and P. Godfrey-Smith. 2009. Generalization of the Price equation for evolutionary change. Evolution 63:531-536. (PDF)


  • Prado, F., and B. Kerr. 2008. The evolution of restraint in bacterial biofilms under nontransitive competition. Evolution 62: 538-548. (PDF)
  • Nahum, J., and B. Kerr. 2008. Optimal foraging: A bird in the hand released. Current Biology 18: R385-R386. (PDF)
  • Kerr, B., J. D. West, B. J. M. Bohannan. 2008. "Bacteriophages: Models for exploring basic principles of ecology." Bacteriophage ecology: Population growth, evolution, and impact of bacterial viruses. Ed. S. T. Abedon. Cambridge, U.K.: University Press, 31-63. (Link)
  • Bergstrom, C., B. Kerr, and M. Lachmann. 2008. "Building trust by wasting time." Moral Markets. Ed. P. J. Zak. Oxfordshire: Princeton University Press, 142-153. (Link)


  • Kerr, B. 2007. Niche construction and cognitive evolution. Biological Theory 2: 250-262. (PDF)
  • Kerr, B. 2007. "The ecological and evolutionary dynamics of model bacteriocin communities." Bacteriocins: Ecology and Evolution. Ed. M.A. Riley and M. A. Chavan. Heidelberg: Springer, 111-134. (PDF)


  • Kerr, B., C. Neuhauser, B. J. M. Bohannan, and A. M. Dean. 2006. Local migration promotes competitive restraint in a host-pathogen ‘tragedy of the commons’. Nature 442: 75-78. (PDF) (Supplemental Material) (News Item)


  • Stephens, D. W., B. Kerr, and E. Fernández-Juricic. 2004. Impulsiveness without discounting: The ecological rationality hypothesis. Proceedings of the Royal Society, Series B 271: 2459-2465 (PDF) (Supplemental Material)
  • Kerr, B. 2004. The Caucas-race of the Dodo: A review of Genetic and Cultural Evolution of Cooperation. Biology and Philosophy 19: 781-799. (PDF)
  • Fernández-Juricic, E., B. Kerr, P. A. Bednekoff, and D. W. Stephens. 2004. When are two heads better than one? Visual perception and information transfer affect the value of vigilance coordination in foraging groups. Behavioral Ecology 15: 898-906. (PDF)
  • Jessup, C., R. Kassen, S. E. Forde, B. Kerr, A. Buckling, P. B. Rainey, and B. J. M. Bohannan. 2004. Asking big questions of small worlds: Microbial model systems in ecological research. Trends in Ecology and Evolution 19: 189-197. (PDF)
  • Kerr, B., P. Godfrey-Smith and M. W. Feldman. 2004. What is altruism? Trends in Ecology and Evolution 19: 135-140. (PDF)


  • Kerr, B., and M. W. Feldman. 2003. Carving the cognitive niche: Optimal learning strategies in homogeneous and heterogeneous environments. Journal of Theoretical Biology 220: 169-188. (PDF)


  • Kerr, B., M. A. Riley, M. W. Feldman, and B. J. M. Bohannan. 2002. Local dispersal promotes biodiversity in a real-life game of rock-paper-scissors. Nature 418 (11): 171-174. (PDF) (Commentary)
  • Bohannan, B. J. M., B. Kerr, C. Jessup, J. Hughes, and G. Sandvik. 2002. Trade-offs and coexistence in microbial microcosms. Antonie von Leeuvenhoek 81: 107-115. (PDF)
  • Kerr, B. and P. Godfrey-Smith. 2002. On individualist and multi-level perspectives on selection in structured populations. Biology and Philosophy 17: 477-517. (PDF)
  • Kerr, B. and P. Godfrey-Smith. 2002. On Price's equation and average fitness. Biology and Philosophy 17: 551-565. (PDF) (Commentary)
  • Godfrey-Smith, P. and B. Kerr. 2002. Group fitness and multi-level selection: Replies to commentaries. Biology and Philosophy 17: 539-550. (PDF)
  • Schwilk, D. W. and B. Kerr. 2002. Genetic niche-hiking: An alternative explanation for the evolution of flammability. Oikos 99: 431-442. (PDF)


  • Aoki, K., M. W. Feldman, and B. Kerr. 2001. Models of sexual selection on a quantitative genetic trait when preference is acquired by sexual imprinting. Evolution 55: 25-32. (PDF)


  • Kerr, B., D. W. Schwilk, A. Bergman, and M. W. Feldman. 1999. Rekindling an old flame: A haploid model for the evolution and impact of flammability in resprouting plants. Evolutionary Ecology Research 1: 807-833. (PDF)